Annual report pursuant to Section 13 and 15(d)

Revenue - Collaboration Agreement - The GSK Collaboration and License Agreement (Details)

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Revenue - Collaboration Agreement - The GSK Collaboration and License Agreement (Details) - USD ($)
1 Months Ended 12 Months Ended
Jun. 30, 2021
Dec. 31, 2021
Dec. 31, 2019
Collaboration and License Agreement      
Amount of the transaction price that is allocated to performance obligations that are unsatisfied or partially satisfied   $ 7,424,000  
GSK Collaboration And License Agreement      
Collaboration and License Agreement      
Milestone payments received $ 4,200,000    
Amount of the transaction price that is allocated to performance obligations that are unsatisfied or partially satisfied   $ 803,000  
GlaxoSmithKline Intellectual Property Development Ltd      
Collaboration and License Agreement      
Amount of development milestone achieved     $ 3,200,000